Biology: Class 11 — Chapter 5: Morphology of Flowering Plants

Introduction to Plant Morphology

Angiosperms exhibit a wide range of structural diversity, yet they are characterized by the presence of roots, stems, leaves, flowers, and fruits. The study of these external features is known as morphology.

5.1 The Root

In majority of dicotyledonous plants, the direct elongation of the radicle leads to the formation of the primary root which grows inside the soil.

Types of Root Systems

  • Tap Root System: Consists of the primary root and its lateral branches (secondary, tertiary roots). Seen in dicotyledonous plants like the mustard plant.
  • Fibrous Root System: The primary root is short-lived and replaced by a large number of roots originating from the base of the stem. Seen in monocotyledonous plants like wheat.
  • Adventitious Roots: Roots that arise from parts of the plant other than the radicle. Examples include grass, Monstera, and the banyan tree.

Regions of the Root

  1. Root Cap: A thimble-like structure covering the apex of the root, protecting its tender tip as it makes its way through the soil.
  2. Region of Meristematic Activity: Located a few millimeters above the root cap. Cells are small, thin-walled, with dense protoplasm, and divide repeatedly.
  3. Region of Elongation: Cells undergo rapid elongation and enlargement, responsible for the growth of the root in length.
  4. Region of Maturation: Cells gradually differentiate and mature. Epidermal cells in this region form very fine, delicate, thread-like structures called root hairs, which absorb water and minerals from the soil.
5.2 The Stem

The stem is the ascending part of the axis bearing branches, leaves, flowers, and fruits. It develops from the plumule of the embryo of a germinating seed.

5.3 The Leaf

The leaf is a lateral, generally flattened structure borne on the stem. It develops at the node and bears an axillary bud in its axil, which later develops into a branch.

Parts of a Leaf

  • Leaf Base: Attaches the leaf to the stem. May bear two lateral small leaf-like structures called stipules. In leguminous plants, the leaf base may become swollen, called the pulvinus.
  • Petiole: Helps hold the leaf blade to light and allows it to flutter in the wind, cooling the leaf.
  • Lamina (Leaf Blade): The green expanded part with veins and veinlets. The middle prominent vein is the midrib.

Venation

The arrangement of veins and veinlets in the lamina of a leaf is called venation.

  • Reticulate Venation: Veinlets form a network; characteristic of most dicotyledonous plants.
  • Parallel Venation: Veins run parallel to each other; characteristic of most monocotyledonous plants.

Types of Leaves

  • Simple Leaf: Lamina is entire or, when incised, incisions do not touch the midrib.
  • Compound Leaf: Incisions of the lamina reach the midrib, breaking it into leaflets. A bud is present in the axil of the petiole but not in the axil of leaflets.
  • Pinnately Compound Leaf: Leaflets are present on a common axis, the rachis (e.g., neem).
  • Palmately Compound Leaf: Leaflets are attached at a common point at the tip of the petiole (e.g., silk cotton).

Phyllotaxy

The pattern of arrangement of leaves on the stem or branch:

  • Alternate: A single leaf arises at each node in an alternate manner (e.g., china rose, mustard, sunflower).
  • Opposite: A pair of leaves arise at each node and lie opposite to each other (e.g., Calotropis, guava).
  • Whorled: More than two leaves arise at a node and form a whorl (e.g., Alstonia).
5.4 The Inflorescence

The arrangement of flowers on the floral axis is termed as inflorescence.

5.5 The Flower

The flower is the reproductive unit in angiosperms, meant for sexual reproduction.

Symmetry of Flowers

  • Actinomorphic (Radial Symmetry): Can be divided into two equal radial halves in any radial plane passing through the center (e.g., mustard, datura, chilli).
  • Zygomorphic (Bilateral Symmetry): Can be divided into two similar halves only in one particular vertical plane (e.g., pea, gulmohur, bean, Cassia).
  • Asymmetric (Irregular): Cannot be divided into two similar halves by any vertical plane passing through the center (e.g., canna).

Parts of a Flower

A typical flower has four whorls arranged on the swollen end of the stalk (thalamus or receptacle):

  1. Calyx: Outermost whorl composed of sepals. Can be gamosepalous (sepals united) or polysepalous (sepals free).
  2. Corolla: Composed of petals. Can be gamopetalous (petals united) or polypetalous (petals free).
  3. Androecium: Composed of stamens (male reproductive organ). Each stamen consists of a filament and a bilobed anther containing pollen-sacs.
  4. Gynoecium: Female reproductive part composed of one or more carpels (stigma, style, and ovary).

Aestivation

The mode of arrangement of sepals or petals in a floral bud with respect to other members of the same whorl:

  • Valvate: Sepals or petals just touch one another at the margin without overlapping (e.g., Calotropis).
  • Twisted: One margin of the appendage overlaps that of the next one (e.g., china rose, lady's finger, cotton).
  • Imbricate: Margins overlap but not in any particular direction (e.g., Cassia, gulmohur).
  • Vexillary (Papilionaceous): Five petals: largest (standard) overlaps two lateral petals (wings), which overlap the two smallest anterior petals (keel) (e.g., pea, bean).

Placentation

The arrangement of ovules within the ovary is known as placentation:

  • Marginal: Placenta forms a ridge along the ventral suture of the ovary (e.g., pea).
  • Axile: Placenta is axial and ovules are attached to it in a multilocular ovary (e.g., china rose, tomato, lemon).
  • Parietal: Ovules develop on the inner wall of the ovary (e.g., mustard, Argemone).
  • Free Central: Ovules are borne on the central axis and septa are absent (e.g., Dianthus, Primrose).
  • Basal: Placenta develops at the base of the ovary and a single ovule is attached to it (e.g., sunflower, marigold).
5.6 The Fruit

The fruit is a mature or ripened ovary developed after fertilization. If formed without fertilization, it is called a parthenocarpic fruit.

5.7 The Seed

The ovules after fertilization develop into seeds. A seed is made up of a seed coat and an embryo.

Structure of a Dicotyledonous Seed

  • Seed Coat: Outer testa and inner tegmen.
  • Hilum: A scar on the seed coat where the seed was attached to the fruit.
  • Micropyle: A small pore above the hilum.
  • Embryo: Consists of an embryonal axis and two cotyledons. The ends of the axis are the radicle and the plumule.
  • Endospermic Seeds: Castor (food storing tissue present).
  • Non-endospermic Seeds: Bean, gram, pea (endosperm absent in mature seeds).

Structure of a Monocotyledonous Seed

  • Generally endospermic, but some (like orchids) are non-endospermic.
  • Aleurone Layer: A proteinous layer separating the embryo from the endosperm.
  • Scutellum: The single large, shield-shaped cotyledon.
  • The plumule and radicle are enclosed in protective sheaths called coleoptile and coleorhiza, respectively.
5.8 & 5.9 Semi-Technical Description & Solanaceae

Flowering plants are described using standard symbols in a floral formula:

Family Solanaceae (Potato Family)

  • Vegetative Characters: Mostly herbs, shrubs, and rarely small trees. Leaves are alternate, simple, exstipulate with reticulate venation. Underground stem in potato (Solanum tuberosum).
  • Floral Characters:
    • Inflorescence: Solitary, axillary, or cymose.
    • Flower: Bisexual, actinomorphic.
    • Calyx: Sepals 5, united, persistent, valvate.
    • Corolla: Petals 5, united, valvate.
    • Androecium: Stamens 5, epipetalous.
    • Gynoecium: Bicarpellary, syncarpous, ovary superior, bilocular, placenta swollen with many ovules, axile placentation.
    • Fruits: Berry or capsule.
    • Seeds: Many, endospermous.
  • Floral Formula: $$\oplus \ \Phi \ K_{(5)} \ \overbrace{C_{(5)} \ A_5} \ \underline{G}_{(2)}$$
  • Economic Importance: Source of food (tomato, brinjal, potato), spice (chilli), medicine (belladonna, ashwagandha), fumigatory (tobacco), and ornamentals (petunia).